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Penelope is a mouse from Holland who led a double-life as an infamous pilot known as the Black Baron. She was the founder and reigning champion of the ACES dogfighting competition. She was later unmasked by the Cooper Gangafter which she ed them as their RC specialist up to and during their mission to access the Cooper Vault. During this time, she developed a relationship with Bentley. After the Cooper Vault job, they both began constructing a time machine. During which, however, Penelope disappeared and ed forces with Cyrille Le Paradox 's gang in an attempt to sever all of Bentley's ties to the Cooper Gang and unlock what she believed to be his full potential, which was implied to be world conquest.
Penelope is a technological expert who specializes in RC helicopters and cars. Bentley gets into contact with her since the gang needs an RC specialist for breaking into the Cooper vault. Penelope agreed to them if they defeated her boss the Black Baron in the aerial dogfights competition. It turns out however that Penelope was the Black Baron since she didn't meet the age requirements.
She was initially a bit turned off by Bentley when they met since they posted fake images online and got a crush on Sly. This started to make Bentley a bit jealous. After Penelope got kidnapped by the pirate LeFwee, she realized she did have feelings for Bentley and beat LeFwee in a duel and started dating Bentley.
After the events of Sly 3, she stayed with Bentley and built a whole new lab from scratch around the location of the Cooper Vault. In Thieves in Time, she and Bentley are working on a top secret project the time machinewhen she mysteriously vanishes without a trace. Bentley looks everywhere, but finds nothing.
She does not appear until Sly and the gang visit Medieval England. It is revealed that she sided has with Le Paradox and has assumed the identity of the Black Knight, which is a robot she has built, and has built numerous mechanical guards and the Moat Monster. Her involvement is discovered when Bentley follows the Black Knight into the blacksmith shop and sees her exit the robot.
It is also revealed that she stole the plans for the time machine from Bentley and gave them to Le Paradox, which is how he developed his own time machine. She later confronts Sly in a giant mech, but Sly manages to destroy it.
When she attempts to kill him, she is stopped by Bentley. When asked why, she says she did it for him, to make him see what he is capable of if they ed forces and created advanced technology together, instead of continuing to work with Sly. When Bentley says she is just Le Paradox's puppet, she becomes angry and attacks him.
She is defeated and sent to Europe's highest security prison, where she promptly escapes from. Her whereabouts are unknown, but she has been sending Bentley some mysterious postcards with her inia on them.
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